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=== Chapter 1 ===
=== Chapter 1 ===
Cerval, remembering the smell of Japari Buns thanks to Serval's memory, happens upon Caracal and attacks her to steal some buns. This action causes a misunderstanding and drives a temporary rift between Serval and Caracal, with the latter mistaking Cerval's actions for those of the real Friend. In order to prove Serval's innocence, the player's party pursues Cerval and -- unbeknownst to them, with the slight aid of Silver Fox, who is collecting data on her -- manages to catch up with her for a confrontation. Not wanting to be bothered, Cerval attacks them but is defeated, uttering a cryptic line about "the Queen". Serval, not knowing what she's referring to, asks for her name, but Cerval does not seem to understand and flees the scene. The party, now including Caracal, makes the decision to pursue her out of concern that she'll end up falsely incriminating Serval of further wrongdoing.
Cerval, remembering the smell of Japari Buns thanks to Serval's memory, happens upon [[Caracal/Nexon Game|Caracal]] and attacks her to steal some buns. This action causes a misunderstanding and drives a temporary rift between Serval and Caracal, with the latter mistaking Cerval's actions for those of the real Friend. In order to prove Serval's innocence, the player's party pursues Cerval and -- unbeknownst to them, with the slight aid of [[Silver Fox/Nexon Game|Silver Fox]], who is collecting data on her -- manages to catch up with her for a confrontation. Not wanting to be bothered, Cerval attacks them but is defeated, uttering a cryptic line about "the Queen". Serval, not knowing what she's referring to, asks for her name, but Cerval does not seem to understand and flees the scene. The party, now including Caracal, makes the decision to pursue her out of concern that she'll end up falsely incriminating Serval of further wrongdoing.


=== Chapter 2 ===
=== Chapter 2 ===
Cerval does not directly appear during Chapter 2, but the player's party receives information from Northern White-Faced Owl and her assistant Eurasian Eagle-Owl suggesting that a Cellien matching Cerval's description had been sighted heading towards the Desert Area. After their business in the forest is complete, the protagonists pursue her there.
Cerval does not directly appear during Chapter 2, but the player's party receives information from [[Northern White-Faced Owl/Nexon Game|Northern White-Faced Owl]] and her assistant [[Eurasian Eagle-Owl/Nexon Game|Eurasian Eagle-Owl]] suggesting that a Cellien matching Cerval's description had been sighted heading towards the Desert Area. After their business in the forest is complete, the protagonists pursue her there.


=== Chapter 3 ===
=== Chapter 3 ===
The Thomson's Gazelle nicknamed Lulu happens upon Cerval when she has been surrounded by Celliens and, not knowing what kind of creature she is, calls out to her for help. Cerval responds, but doesn't understand what "help" means. Lulu attempts to explain, but Cerval misunderstands further, calling forth even more Celliens and making the situation even more dire. Erroneously believing that she's "lent her power" as requested, she exits the scene promptly.
The [[Thomson's Gazelle/Nexon Game|Thomson's Gazelle]] nicknamed Lulu happens upon Cerval when she has been surrounded by Celliens and, not knowing what kind of creature she is, calls out to her for help. Cerval responds, but doesn't understand what "help" means. Lulu attempts to explain, but Cerval misunderstands further, calling forth even more Celliens and making the situation even more dire. Erroneously believing that she's "lent her power" as requested, she exits the scene promptly.


At the end of the chapter, when Lulu is safely reunited with her friend Rabi-Rabi the Arabian Oryx and the rest of the party, the subject of the Serval lookalike is brought up again amongst the heroes. Lulu, remembering the girl who tried to help her, mentions that she seemed to head off towards the Waterfront Area, and thus they head in that direction in pursuit of Cerval.
At the end of the chapter, when Lulu is safely reunited with her friend Rabi-Rabi the [[Arabian Oryx/Nexon Game|Arabian Oryx]] and the rest of the party, the subject of the Serval lookalike is brought up again amongst the heroes. Lulu, remembering the girl who tried to help her, mentions that she seemed to head off towards the Waterfront Area, and thus they head in that direction in pursuit of Cerval.


=== Chapter 4 ===
=== Chapter 4 ===
During this chapter, a mysterious disease has been spreading in the Waterfront Area, which White Rhinoceros and Black Rhinoceros -- after initially believing Serval to be the culprit -- suspect the "Fake Serval" to be behind. Together with the protagonist's party, they investigate the area in hopes of getting to the bottom of the outbreak.
During this chapter, a mysterious disease has been spreading in the Waterfront Area, which [[White Rhinoceros/Nexon Game|White Rhinoceros]] and [[Black Rhinoceros/Nexon Game|Black Rhinoceros]] -- after initially believing Serval to be the culprit -- suspect the "Fake Serval" to be behind. Together with the protagonist's party, they investigate the area in hopes of getting to the bottom of the outbreak.


During a short break where Hippopotamus offers the group some Japari Buns, Cerval (lured by her favorite food) finds the party herself, to their surprise. When she finds they've already finished their meal by the time she gets there, though, she leaves promptly, but the group gives chase, accumulating in another direct confrontation between the player's party and Cerval. To Serval's surprise, Cerval is actually able to understand her words despite having contracted the disease ("Gaogao Disease", which makes it impossible for Friends to speak in anything other than animal noises by stealing away the Sparkle of Words from them). The confrontation with Cerval accumulates in another battle, where she is yet again defeated and this time bound by a rope so she cannot escape.
During a short break where [[Hippopotamus/Nexon Game|Hippopotamus]] offers the group some Japari Buns, Cerval (lured by her favorite food) finds the party herself, to their surprise. When she finds they've already finished their meal by the time she gets there, though, she leaves promptly, but the group gives chase, accumulating in another direct confrontation between the player's party and Cerval. To Serval's surprise, Cerval is actually able to understand her words despite having contracted the disease ("Gaogao Disease", which makes it impossible for Friends to speak in anything other than animal noises by stealing away the Sparkle of Words from them). The confrontation with Cerval accumulates in another battle, where she is yet again defeated and this time bound by a rope so she cannot escape.


The captured Cerval accompanies the group for a leg of the journey and exhibits subtly different behavior during this time, even expressing interest in seeing a scenic waterfall they pass by during their travel. However, once they reach their destination -- the water regulation facility called Undine -- she suddenly reverts to her more familiar aloofness, escaping from the ropes she'd been bound in with ease and disappearing again. Interestingly, she calls out to Serval once before leaving, bidding her farewell in a warmer tone before becoming cold once more and fleeing the scene. Serval laments (although no one can understand her due to the Gaogao Disease) that she still hasn't given her a proper name.
The captured Cerval accompanies the group for a leg of the journey and exhibits subtly different behavior during this time, even expressing interest in seeing a scenic waterfall they pass by during their travel. However, once they reach their destination -- the water regulation facility called Undine -- she suddenly reverts to her more familiar aloofness, escaping from the ropes she'd been bound in with ease and disappearing again. Interestingly, she calls out to Serval once before leaving, bidding her farewell in a warmer tone before becoming cold once more and fleeing the scene. Serval laments (although no one can understand her due to the Gaogao Disease) that she still hasn't given her a proper name.
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Eventually, when Silver Fox's memories return to her and she explains that she was sent by Oinari-sama to watch over the protagonist, Cerval comments to herself that the mention of that name makes her chest feel uneasy. It's not long afterwards that, during a conversation about the origin of the protagonist's special amulet, Cerval suddenly flees the scene and reverts to her cold demeanour, which prompts the party to pursue her. When they catch up to her they see her, with her body glowing like before, raising her hand to a giant boss Cellien to power it up. Like before, she mentions in her trance-like state that she's doing it "for the sake of the Queen". Serval, determined to protect her friend and not believing that these actions reflect the "real" Cerval, charges boldly into battle against the giant Cellien to save her.
Eventually, when Silver Fox's memories return to her and she explains that she was sent by Oinari-sama to watch over the protagonist, Cerval comments to herself that the mention of that name makes her chest feel uneasy. It's not long afterwards that, during a conversation about the origin of the protagonist's special amulet, Cerval suddenly flees the scene and reverts to her cold demeanour, which prompts the party to pursue her. When they catch up to her they see her, with her body glowing like before, raising her hand to a giant boss Cellien to power it up. Like before, she mentions in her trance-like state that she's doing it "for the sake of the Queen". Serval, determined to protect her friend and not believing that these actions reflect the "real" Cerval, charges boldly into battle against the giant Cellien to save her.


After emerging victorious, Cerval appears to come back to her senses temporarily, but falls in and out of her trance-like state and leaves the area in order to achieve her stated goal of returning to the Queen's side. Before leaving, she nearly calls Serval her "friend", but refers her as a "clumsy troublemaker" instead, to the Friend's comical chagrin. After the party decides (like always) to pursue her to the next region, Fennec mentions to them that she and Arai-san are planning on pursuing Cerval as well, along with a cryptic implication that she is "the enemy" after all. This does not seem to deter Serval's spirits, though, and she and her friends boldly pursue her through the Hotoku Region.
After emerging victorious, Cerval appears to come back to her senses temporarily, but falls in and out of her trance-like state and leaves the area in order to achieve her stated goal of returning to the Queen's side. Before leaving, she nearly calls Serval her "friend", but refers her as a "clumsy troublemaker" instead, to the Friend's comical chagrin. After the party decides (like always) to pursue her to the next region, [[Fennec Fox/Nexon Game|Fennec]] mentions to them that she and [[Raccoon/Nexon Game|Arai-san]] are planning on pursuing Cerval as well, along with a cryptic implication that she is "the enemy" after all. This does not seem to deter Serval's spirits, though, and she and her friends boldly pursue her through the Hotoku Region.


=== Chapter 6 ===
=== Chapter 6 ===
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=== Chapter 7 ===
=== Chapter 7 ===
Whilst helping Margay shoot her movie, the party happens upon an ominous "ghost ship" on the water in the vicinity of their film set. Upon examination, it's revealed that Cerval is, in fact, at its helm, and its crew is comprised entirely of black Celliens. Rather than attacking the protagonist and their group, however, "Captain Cerval" -- which she introduces herself as -- informs the protagonists that they are playing pirates, going off of Margay's script (which the Celliens stole to begin with). Cerval and her crew of Celliens receive the compliments and admiration of the film-making Friends for their impeccable coordination and acting skills, and Margay makes the bold decision to invite them to help her finish the movie. Captain Cerval (and her Cellien crew) readily accepts the offer, expressing that she wants to be in a movie, and their collaboration begins.
Whilst helping [[Margay/Nexon Game|Margay]] shoot her movie, the party happens upon an ominous "ghost ship" on the water in the vicinity of their film set. Upon examination, it's revealed that Cerval is, in fact, at its helm, and its crew is comprised entirely of black Celliens. Rather than attacking the protagonist and their group, however, "Captain Cerval" -- which she introduces herself as -- informs the protagonists that they are playing pirates, going off of Margay's script (which the Celliens stole to begin with). Cerval and her crew of Celliens receive the compliments and admiration of the film-making Friends for their impeccable coordination and acting skills, and Margay makes the bold decision to invite them to help her finish the movie. Captain Cerval (and her Cellien crew) readily accepts the offer, expressing that she wants to be in a movie, and their collaboration begins.


In the film story, Cerval is cast as (Captain) Serval's former friend, whose mind is manipulated by the ghost ship in order to force her to protect its treasure. The scene opens with Serval boarding the ghost ship and trying in vain to snap the Cellien back to her senses (not unlike some of their past interactions, which were not acting). The possessed Captain Cerval commands her underlings to attack Serval and her crew, and due to the Cellien crew's incredible coordination they are able to easily overwhelm the heroes. The battle rages on, however, and with the help of multiple dolphin Friends, Captain Serval is able to turn the tide in her favor.
In the film story, Cerval is cast as (Captain) Serval's former friend, whose mind is manipulated by the ghost ship in order to force her to protect its treasure. The scene opens with Serval boarding the ghost ship and trying in vain to snap the Cellien back to her senses (not unlike some of their past interactions, which were not acting). The possessed Captain Cerval commands her underlings to attack Serval and her crew, and due to the Cellien crew's incredible coordination they are able to easily overwhelm the heroes. The battle rages on, however, and with the help of multiple dolphin Friends, Captain Serval is able to turn the tide in her favor.
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=== Chapter 8 ===
=== Chapter 8 ===
The party follows Cerval's path to the Riukiu Region, where they meet Shisa Lefty and Shisa Right. The Shisas inform them that she has fallen onto one of the region's islands, and since then, large numbers of black Celliens (much like the ones who served as her "crew" in the previous chapter) have gathered around the island, as if to protect her. Wasting no time, the Friends of Riukiu have already begun preparations so that the protagonists can break through the Cellien barricade and reach her again. They stay the night in Riukiu as further information is gathered on her situation.
The party follows Cerval's path to the Riukiu Region, where they meet [[Shisa Lefty/Nexon Game|Shisa Lefty]] and [[Shisa Right/Nexon Game|Shisa Right]]. The Shisas inform them that she has fallen onto one of the region's islands, and since then, large numbers of black Celliens (much like the ones who served as her "crew" in the previous chapter) have gathered around the island, as if to protect her. Wasting no time, the Friends of Riukiu have already begun preparations so that the protagonists can break through the Cellien barricade and reach her again. They stay the night in Riukiu as further information is gathered on her situation.


When preparations have been made for an invasion of the island, the Shisas also take the time to inform Serval and her friends of some key details regarding the Cellien Queen's plan -- and how Cerval figures into it. They tell them that the Queen, knowing that Cerval possesses the "special something" necessary for Kemo Harmony, plans to invoke a Cellien equivalent called "Cell Harmony" instead using her power. They point out that in the time Cerval has been active, she's powered up other Celliens in various ways, not the least of which being the new black Celliens, which are revealed to be evolved from regular specimens. The Queen's plans, however, don't end there -- it intends to spread these properties to all Celliens in the Park, accumulating in something called "Ultimate Cell Harmony", wherein all Celliens would evolve monstrously together. If that happened, the Shisas propose, they'd be able to steal all the Sparkle in Japari Park without opposition and take over this world with ease. Expectedly, the party is unnerved by this cataclysmic prospect, and their resolve is strengthened to retrieve the "special something" that Cerval stole to prevent it all from happening.
When preparations have been made for an invasion of the island, the Shisas also take the time to inform Serval and her friends of some key details regarding the Cellien Queen's plan -- and how Cerval figures into it. They tell them that the Queen, knowing that Cerval possesses the "special something" necessary for Kemo Harmony, plans to invoke a Cellien equivalent called "Cell Harmony" instead using her power. They point out that in the time Cerval has been active, she's powered up other Celliens in various ways, not the least of which being the new black Celliens, which are revealed to be evolved from regular specimens. The Queen's plans, however, don't end there -- it intends to spread these properties to all Celliens in the Park, accumulating in something called "Ultimate Cell Harmony", wherein all Celliens would evolve monstrously together. If that happened, the Shisas propose, they'd be able to steal all the Sparkle in Japari Park without opposition and take over this world with ease. Expectedly, the party is unnerved by this cataclysmic prospect, and their resolve is strengthened to retrieve the "special something" that Cerval stole to prevent it all from happening.
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As over-the-top as this display is, it touches Cerval's heart deeply. She calls out repeatedly to the collapsed Serval, and her body begins to shine brightly, to the surprise of the (still conscious) party members, who wonder what it could mean. The Cellien Queen herself confirms for everyone that Cerval, a Cellien, has somehow produced a Sparkle of her own, wondering aloud how this could possibly have happened.
As over-the-top as this display is, it touches Cerval's heart deeply. She calls out repeatedly to the collapsed Serval, and her body begins to shine brightly, to the surprise of the (still conscious) party members, who wonder what it could mean. The Cellien Queen herself confirms for everyone that Cerval, a Cellien, has somehow produced a Sparkle of her own, wondering aloud how this could possibly have happened.


Cerval -- who has now gained a new and upgraded form as a Cellien-gone-Friend -- confidently informs the Queen that she's not a Cellien any more, and that she'll no longer be listening to her commands. The Cellien Queen, seeing no more use for her, orders her underlings to "dispose of the mutant". Cerval is undeterred, joining the heroes in their battle against the Cellien Queen's minions and vowing to do her best for everyone's sake, including that of "Serval, who died". (She is quickly informed by the other party members that Serval is, in fact, still alive and just ate too much salt at once.) It is at this point that Cerval becomes useable in gameplay as a regular Friend. Serval regains consciousness, and the heroes -- Cerval included -- take the fight directly to the Cellien Queen. It doesn't come easily, but at long last they triumph over the monstrous final boss of the main plot, freeing Cerval once and for all from the proverbial chains which had bound her the entire plot prior. And just as Serval hoped, it was done without ending her life or the friendship the lookalike pair fostered over the course of the story.
Cerval -- who has now gained a new and upgraded form as a Cellien-gone-Friend -- confidently informs the Queen that she's not a Cellien any more, and that she'll no longer be listening to her commands. The Cellien Queen, seeing no more use for her, orders her underlings to "dispose of the mutant". Cerval is undeterred, joining the heroes in their battle against the Cellien Queen's minions and vowing to do her best for everyone's sake, including that of "Serval, who died". (She is quickly informed by the other party members that Serval is, in fact, still alive and just ate too much salt at once.) It is at this point that Cerval becomes usable in gameplay as a regular Friend. Serval regains consciousness, and the heroes -- Cerval included -- take the fight directly to the Cellien Queen. It doesn't come easily, but at long last they triumph over the monstrous final boss of the main plot, freeing Cerval once and for all from the proverbial chains which had bound her the entire plot prior. And just as Serval hoped, it was done without ending her life or the friendship the lookalike pair fostered over the course of the story.


After the Cellien Queen's defeat, peace returns to Japari Park, and Cerval is able to freely enjoy herself with the friends she made throughout the adventure at long last. When Mirai suggests that the everyone take another trip around the Park, this time for leisure and not to chase Cerval around, the former Cellien happily agrees to accompanying them on the adventure.
After the Cellien Queen's defeat, peace returns to Japari Park, and Cerval is able to freely enjoy herself with the friends she made throughout the adventure at long last. When Mirai suggests that the everyone take another trip around the Park, this time for leisure and not to chase Cerval around, the former Cellien happily agrees to accompanying them on the adventure.